I. Van-seuningen and E. Leteurtre, Mucin expression and regulation in the gastro-intestinal tract. Actors of carcinogenesis? The Epithelial Mucins: Structure/Function, Roles in Cancer and Inflammatory Diseases, pp.183-210, 2008.

M. A. Mcguckin, S. K. Linden, P. Sutton, and T. H. Florin, Mucin dynamics and enteric pathogens, Nature Reviews Microbiology, vol.154, issue.4, pp.265-278
DOI : 10.1038/nrmicro2538

J. Dekker, J. W. Rossen, H. A. Buller, and A. W. Einerhand, The MUC family: an obituary, Trends in Biochemical Sciences, vol.27, issue.3, pp.126-131, 2002.
DOI : 10.1016/S0968-0004(01)02052-7

A. P. Corfield, D. Carroll, N. Myerscough, and C. S. Probert, Mucins in the gastrointestinal tract in health and disease, Frontiers in Bioscience, vol.6, issue.3, pp.1321-1357, 2001.
DOI : 10.2741/A684

P. Pigny, V. Guyonnet-duperat, A. S. Hill, W. S. Pratt, S. Galiegue-zouitina et al., Human Mucin Genes Assigned to 11p15.5: Identification and Organization of a Cluster of Genes, Genomics, vol.38, issue.3, pp.38-340, 1996.
DOI : 10.1006/geno.1996.0637

J. L. Desseyn and A. Laine, Characterization of mouse muc6 and evidence of conservation of the gel-forming mucin gene cluster between human and mouse, Genomics, pp.81-433, 2003.

M. P. Buisine, N. Porchet, and I. Van-seuningen, Mucin expression and regulation during development and cell differentiation The Epithelial Mucins: Structure/Function, Roles in Cancer and Inflammatory Diseases, pp.75-94, 2008.

M. P. Buisine, L. Devisme, T. C. Savidge, C. Gespach, B. Gosselin et al., Mucin gene expression in human embryonic and fetal intestine, Gut, vol.43, issue.4, pp.43-519, 1998.
DOI : 10.1136/gut.43.4.519

M. P. Buisine, L. Devisme, V. Maunoury, E. Deschodt, B. Gosselin et al., Developmental Mucin Gene Expression in the Gastroduodenal Tract and Accessory Digestive Glands. I. Stomach: A Relationship to Gastric Carcinoma, Journal of Histochemistry & Cytochemistry, vol.348, issue.12, pp.48-1657, 2000.
DOI : 10.1177/002215540004801209

I. Van-seuningen, P. Pigny, M. Perrais, N. Porchet, and J. P. Aubert, Transcriptional regulation of the 11p15 mucin genes towards new biological tools in human therapy in inflammatory diseases and cancer, Frontiers in Bioscience, vol.6, issue.3, pp.1216-1234, 2001.
DOI : 10.2741/A675

J. D. Molkentin, The Zinc Finger-containing Transcription Factors GATA-4, -5, and -6: UBIQUITOUSLY EXPRESSED REGULATORS OF TISSUE-SPECIFIC GENE EXPRESSION, Journal of Biological Chemistry, vol.275, issue.50, pp.275-38949, 2000.
DOI : 10.1074/jbc.R000029200

R. K. Montgomery, A. E. Mulberg, and R. J. Grand, Development of the human gastrointestinal tract: Twenty years of progress, Gastroenterology, vol.116, issue.3, pp.116-702, 1999.
DOI : 10.1016/S0016-5085(99)70193-9

C. A. Richmond and D. T. Breault, Regulation of Gene Expression in the Intestinal Epithelium, Prog Mol Biol Transl Sci, pp.96-207, 2010.
DOI : 10.1016/B978-0-12-381280-3.00009-9

P. G. Traber and D. G. Silberg, Intestine-Specific Gene Transcription, Annual Review of Physiology, vol.58, issue.1, pp.275-297, 1996.
DOI : 10.1146/annurev.ph.58.030196.001423

P. Traber and G. D. Wu, Gastrointestinal cancers: Biology, Diagnosis, and Therapy, Lippincott-Raven publishers, 1995.

S. A. Duncan, K. Manova, W. S. Chen, P. Hoodless, D. C. Weinstein et al., Expression of transcription factor HNF-4 in the extraembryonic endoderm, gut, and nephrogenic tissue of the developing mouse embryo: HNF-4 is a marker for primary endoderm in the implanting blastocyst., Proceedings of the National Academy of Sciences, vol.91, issue.16, pp.91-7598, 1994.
DOI : 10.1073/pnas.91.16.7598

S. Cereghini, Liver-enriched transcription factors and hepatocyte differentiation, Faseb J, vol.10, pp.267-282, 1996.

R. Hromas and R. Costa, The hepatocyte nuclear factor-3/forkhead transcription regulatory family in development, inflammation, and neoplasia, Critical Reviews in Oncology/Hematology, vol.20, issue.1-2, pp.129-140, 1995.
DOI : 10.1016/1040-8428(94)00151-I

K. H. Kaestner, H. Hiemisch, B. Luckow, and G. Schutz, The HNF-3 Gene Family of Transcription Factors in Mice: Gene Structure, cDNA Sequence, and mRNA Distribution, Genomics, vol.20, issue.3, pp.20-377, 1994.
DOI : 10.1006/geno.1994.1191

V. Gouyer, F. Gottrand, and J. L. Desseyn, The Extraordinarily Complex but Highly Structured Organization of Intestinal Mucus-Gel Unveiled in Multicolor Images, PLoS ONE, vol.49, issue.4, p.18761, 2011.
DOI : 10.1371/journal.pone.0018761.g002

M. G. Roy, M. Rahmani, J. R. Hernandez, S. N. Alexander, C. Ehre et al., Mucin Production during Prenatal and Postnatal Murine Lung Development, American Journal of Respiratory Cell and Molecular Biology, vol.44, issue.6, pp.44-755, 2011.
DOI : 10.1165/rcmb.2010-0020RC
URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3135838

N. Jonckheere, M. Van-der-sluis, A. Velghe, M. P. Buisine, M. Sutmuller et al., Van Seuningen, Transcriptional activation of the murine Muc5ac mucin gene in epithelial cancer cells by TGF-beta/Smad4 signalling pathway is potentiated by Sp1, Biochem J, pp.377-797, 2004.

I. B. Renes, J. A. Boshuizen, D. J. Van-nispen, N. P. Bulsing, H. A. Buller et al., Alterations in Muc2 biosynthesis and secretion during dextran sulfate sodium-induced colitis, American Journal of Physiology - Gastrointestinal and Liver Physiology, vol.282, issue.2, pp.282-382, 2002.
DOI : 10.1152/ajpgi.00229.2001

P. Mesquita, N. Jonckheere, R. Almeida, M. P. Ducourouble, J. Serpa et al., Human MUC2 Mucin Gene Is Transcriptionally Regulated by Cdx Homeodomain Proteins in Gastrointestinal Carcinoma Cell Lines, Journal of Biological Chemistry, vol.278, issue.51, pp.278-51549, 2003.
DOI : 10.1074/jbc.M309019200

M. Van-der-sluis, M. H. Melis, N. Jonckheere, M. P. Ducourouble, H. A. Buller et al., The murine Muc2 mucin gene is transcriptionally regulated by the zinc-finger GATA-4 transcription factor in intestinal cells, Biochemical and Biophysical Research Communications, vol.325, issue.3, pp.325-952, 2004.
DOI : 10.1016/j.bbrc.2004.10.108

M. Perrais, P. Pigny, M. P. Buisine, N. Porchet, J. P. Aubert et al., Van Seuningen-Lempire, Aberrant expression of human mucin gene MUC5B in gastric carcinoma and cancer cells. Identification and regulation of a distal promoter, J Biol Chem, pp.276-15386, 2001.

I. Van-seuningen, J. Ostrowski, X. R. Bustelo, P. R. Sleath, and K. Bomsztyk, The K Protein Domain That Recruits the Interleukin 1-responsive K Protein Kinase Lies Adjacent to a Cluster of c-Src and Vav SH3-binding Sites: IMPLICATIONS THAT K PROTEIN ACTS AS A DOCKING PLATFORM, Journal of Biological Chemistry, vol.270, issue.45, pp.270-26976, 1995.
DOI : 10.1074/jbc.270.45.26976

I. Van-seuningen, M. Perrais, P. Pigny, N. Porchet, and J. P. Aubert, Sequence of the 5???-flanking region and promoter activity of the human mucin gene MUC5B in different phenotypes of colon cancer cells, Biochemical Journal, vol.348, issue.3, pp.675-686, 2000.
DOI : 10.1042/bj3480675

N. Jonckheere, A. Velghe, M. P. Ducourouble, M. C. Copin, I. B. Renes et al., The mouse Muc5b mucin gene is transcriptionally regulated by thyroid transcription factor-1 (TTF-1) and GATA-6 transcription factors, FEBS Journal, vol.402, issue.Pt 1, pp.278-282, 2011.
DOI : 10.1111/j.1742-4658.2010.07945.x
URL : https://hal.archives-ouvertes.fr/inserm-00807850

R. K. Patient and J. D. Mcghee, The GATA family (vertebrates and invertebrates), Current Opinion in Genetics & Development, vol.12, issue.4, pp.416-422, 2002.
DOI : 10.1016/S0959-437X(02)00319-2

C. M. Jacobsen, N. Narita, M. Bielinska, A. J. Syder, J. I. Gordon et al., Genetic Mosaic Analysis Reveals That GATA-4 Is Required for Proper Differentiation of Mouse Gastric Epithelium, Developmental Biology, vol.241, issue.1, pp.241-275, 2002.
DOI : 10.1006/dbio.2001.0424

P. Bossard and K. S. Zaret, GATA transcription factors as potentiators of gut endoderm differentiation, Development, pp.125-4909, 1998.

S. D. Krasinski, H. M. Van-wering, M. R. Tannemaat, and R. J. Grand, Differential activation of intestinal gene promoters: functional interactions between GATA-5 and HNF-1 alpha, Am J Physiol Gastrointest Liver Physiol, pp.281-69, 2001.

F. Boudreau, E. H. Rings, H. M. Van-wering, R. K. Kim, G. P. Swain et al., Hepatocyte nuclear factor-1 alpha, GATA-4, and caudal related homeodomain protein Cdx2 interact functionally to modulate intestinal gene transcription. Implication for the developmental regulation of the sucrase-isomaltase gene, J Biol Chem, pp.277-31909, 2002.

J. N. Rottman and J. I. Gordon, Comparison of the patterns of expression of rat intestinal fatty acid binding protein/human growth hormone fusion genes in cultured intestinal epithelial cell lines and in the gut epithelium of transgenic mice, J Biol Chem, pp.268-11994, 1993.

R. J. Rings, S. D. Grand, and . Krasinski, Physical interaction between GATA-5 and hepatocyte nuclear factor-1alpha results in synergistic activation of the human lactase-phlorizin hydrolase promoter, J Biol Chem, pp.277-27659, 2002.

M. Van-der-sluis, A. Vincent, J. Bouma, A. Korteland-van-male, J. B. Van-goudoever et al., Forkhead box transcription factors Foxa1 and Foxa2 are important regulators of Muc2 mucin expression in intestinal epithelial cells, Biochemical and Biophysical Research Communications, vol.369, issue.4, pp.369-1108, 2008.
DOI : 10.1016/j.bbrc.2008.02.158

J. N. Carraway, I. B. Freund, I. Renes, and . Van-seuningen, The human mucin MUC4 is transcriptionally regulated by caudal-related homeobox, hepatocyte nuclear factors, forkhead box A, and GATA endodermal transcription factors in epithelial cancer cells, J Biol Chem, pp.282-22638, 2007.

E. E. Morrisey, H. S. Ip, M. M. Lu, and M. S. Parmacek, GATA-6: A Zinc Finger Transcription Factor That Is Expressed in Multiple Cell Lineages Derived from Lateral Mesoderm, Developmental Biology, vol.177, issue.1, pp.177-309, 1996.
DOI : 10.1006/dbio.1996.0165

A. C. Laverriere, C. Macneill, C. Mueller, R. E. Poelmann, J. B. Burch et al., GATA-4/5/6, a subfamily of three transcription factors transcribed in developing heart and gut, J Biol Chem, pp.269-23177, 1994.

T. A. Wilson, R. K. Wynn, D. J. Grencis, and . Thornton, Muc5ac: a critical component mediating the rejection of enteric nematodes, J Exp Med, vol.208, pp.893-900, 2011.